Planet Earth +?:
Allo-Hybridizing
Interactions
of
Self-Hybrid
eukaryotic
cell units
and/versus
Self-Hybrid
prokaryotic
cell units.
Part 6.:
DialecticS
of
NATURE
Main
Sequence
Series.
Dear Reader,
The overall ‘dialectic of
Nature’ ontological-categorial progression model includes the dialectical
partial synthesis category, ‘synphysis’ category,
or ‘uniphysis’ category, symbolized by qep, in its 7th model
epoch.
That epoch ‘‘‘contains’’’ the
ancient macro-cosmological hybridization
processes of eukaryote-units-interacting-with-prokaryote-units, still
ongoing today, but only known to occur, so far, on planet Earth.
The epoch 7 model is excerpted,
below, using the NQ
dialectical language’s ‘dyadic Seldon function’, with that
function-form operating upon the ‘Regenerist’ Hypothesis “Dark Energy” ‘spandetron’ units’ «arché»-category, qx, and is, partially [‘¶’], solved by us
as follows –
qx27
= qx128 ¶|-º
qx <+> qc <+> qcx <+> qr <+>
qrx <+> qrc <+> qrcx <+> qa <+>
qax <+> qac <+> qacx <+> qar <+>
qarx <+> qarc <+> qarcx <+> qm <+>
qmx <+> qmc <+> qmcx <+> qmr <+>
qmrx <+> qmrc <+> qmrcx <+> qma <+>
qmax <+> qmac <+> qmacx <+> qmar <+>
qmarx <+> qmarc <+> qmarcx <+> qp <+>
. . . <+>
qep
– with ontological category qep, corresponding to generic ordinal qualifier category q96.
Category qep is partially solved-for
[‘¶|-º’], by us as representing, primarily,
the [cosmo-]ontological category of processes of direct ingestions of prokaryotic living
cell units, qp, by eukaryotic living cell units, qe, as well as their ‘exo-symbiotic
natural formations’, of “mutualist”, that is, of mutually-gainful coalitions
between eukaryotic
living cell units and prokaryotic living
cell units.
“Eukaryotic Living Cells” vs. “Prokaryotic
Living Cells”. Within the
above-written overall ontological, dialectical [i.e., ‘multi-«aufheben»-izing’,
‘multi-meta-unit-izing’] categorial progression, is the sub-progression that
forms the focus of this blog-entry, namely –
qe <x> qp = qp <+> qep.
This component-subprocess,
categorizes and ‘character-izes’ [i.e., ‘algebra-izes’]
– “stands for” – eukaryotic
living cell units/prokaryotic living
cell units inter-actions.
Note: Our research on this hybrid
ontological category has been assisted by AI-researchers.
Some Known Planet-Earth Instantiations of ‘Dialectical
Partial-Synthesis’ Category qep.
This ‘merely-hybrid’
ontological category is exemplary of dialectical synthesis categories in
general in the dialectics of nature.
For its cosmo-ontological model
epoch, model epoch 7,
with a total of 27 or 128 ontological categories possibly extant, category qep [---> q96 is a partial ‘dialectical
synthesis category’ or ‘uni-category’, because it does not combine the subscripts
of all of the ‘self-hybrid categories’ that are extant in model-epoch 7. The epoch 7 ‘full synthesis’ hybrid category/‘uni-category’ is
qepmarcx [---> q127.
The meaning of
category-symbol qep encompasses three
major sub-categories of e versus
p
interactions: (1.) ‘endo-symbiotic’
interactions (2.) predatory
interactions, and what we call (3.) ‘exo-symbiotic interactions, each addressed
separately below’
1. Ancient, Preliminary, Enabling “Auto-Endo-Symbiotic”
Interaction:
qpp à qe.
Approximately 1.5 billion
years ago, an ancient bacterial prokaryotic cell unit – specifically an
alphaproteobacterium unit – was ingested, but not
digested, by a predatory prokaryotic cell unit, likely one related to the Asgard
Archaea:
qp <x> qp =
qp <+> qpp |-º qp <+> qe.
This ‘self-hybridization’ of prokaryotes led
to a lasting “endosymbiotic” relationship, which allowed the ingesting archaean
prokaryotic
cell unit to utilize the bacterial prokaryotic cell unit’s ability to produce ATP
bio-chemical energy more efficiently, i.e., by Oxygen-using respiration,
rather than by anaerobic fermentation, thence powering the subsequent evolution
of modern eukaryotic
cell units.
2. Contemporary Predatory
Interactions Examples.
Single-celled eukaryotes that
eat prokaryotes (bacteria and archaea) are known as “bacterivorous protists”.
They hunt and consume
single-celled prokaryotes through a process called “phagocytosis” (cellular
engulfment), which shapes bacterial populations in almost every ecosystem on
Earth.
In the context of this ontological
sub-category, category-symbol qep connotes ‘qualo-fractal
up-conversions’ of category p or qp biomass into
category e or
qe biomass.
Below is a breakdown of
specific eukaryotic single-cell predator species paired with the specific
prokaryotic prey species they ingest.
1. Tetrahymena
thermophila (Ciliate)
Tetrahymena
thermophila is a highly motile, pear-shaped freshwater ciliate
eukaryote. It uses rows of beating
hair-like structures (cilia) to generate microscopic water currents, sweeping
bacterial cells into its oral groove (cell mouth).
2. Acanthamoeba
castellanii (Amoeba)
Acanthamoeba castellanii is a free-living, amorphous amoebic eukaryotic cell
unit, found ubiquitously in soil and water environments.
It extends flexible,
spine-like cellular extensions called “acanthopodia” to physically trap and
envelop surface-bound bacterial prokaryote units.
- Prokaryote
Prey Species:
- Klebsiella
pneumoniae (A rod-shaped, capsule-forming bacterial prokaryote
units-kind).
- Staphylococcus
aureus (A spherical, Gram-positive cluster bacterial prokaryote
units-kind).
- Legionella
pneumophila (The bacterial prokaryote responsible for
Legionnaires’ disease, which the Acanthamoeba
eukaryotes naturally hunt, though this bacterial prokaryote can sometimes
escape digestion and live inside this eukaryote).
3. Paramecium
caudatum (Ciliate)
Paramecium
caudatum is a slipper-shaped, large eukaryotic single cell unit, widely
found in stagnant freshwater basins. Even a single Paramecium
cell unit is a voracious filter-feeder, capable of consuming up to 5,000
prokaryotes every day.
4. Dictyostelium
discoideum (Social Amoeba/Slime Mold)
While Dictyostelium
discoideum eukaryotes can aggregate into multicellular structures when
starving, it spends its primary life stage as a single-celled amoeba unit, crawling
through forest soil and leaf litter to hunt prokaryotes.
- Prokaryote
Prey Species:
- Klebsiella
aerogenes (The standard target species used to cultivate this
amoeba in laboratory research).
- Salmonella
enterica (A pathogenic bacterial prokaryote unit that this eukaryotic
soil amoeba naturally targets and clears, through phagocytosis).
Summary Table,
Predatory qep Sub-Category Examples.
3. [‘Exo-’]Symbiotic qep Sub-Category
Examples.
Highly complex, mutualistic
coalitions between eukaryotic single cell units and prokaryotic single cell
units are rife on our planet.
In these non-predatory,
non-parasitic associations, the distinct cellular boundaries remain intact, but
the organisms act as a single, coordinated physiological unit (known as a “holobiont”).
These stable coalitions
represent both evolutionary bridges (echoing how mitochondria and chloroplasts
originally evolved) and highly specialized modern ecological teams.
qep, Eukaryote-Prokaryote Coalitions Examples.
|
Eukaryotic Host (Single Eukaryotic
Cell Units)
|
Prokaryotic Partner(s) (Single Prokaryotic Cell Units)
|
Nature of Relationship
|
Biological Function
|
|
Mixotricha
paradoxa
(Parabasalid flagellate)
|
* Treponema mixotrichae (Spirochete)
* Synergitannerella mixotrichae (Rod)
* Endomicrobiellum mixotrichae (Spherical internal bacterium)
|
Ectosymbiosis & Endosymbiosis
(Multi-partner coalition)
|
Locomotion and Digestion: The prokaryotic
spirochetes lash in unison to move the eukaryotic host. The internal bacterial
prokaryotes digest wood and produce energy in place of mitochondria.
|
|
Paulinella
chromatophora
(Filose amoeba)
|
Synechococcus-like
alpha-cyanobacterium
(Termed a "chromatophore")
|
Obligate Endosymbiosis
(Primary plastid in-the-making)
|
Photosynthesis: The eukaryotic host feeds the
bacterial prokaryotes nutrients; the
bacterium has lost 74% of its genome and uses light to supply carbohydrates
to the host.
|
|
Epithemia
turgida
(Diatom alga)
|
Spheroid Body
(UCYN-A related nitrogen-fixing cyanobacteria)
|
Obligate Endosymbiosis
(metabolic coalition)
|
Nitrogen Fixation: The eukaryotic diatom provides
photosynthetic byproducts to the embedded prokaryote, which fixes nitrogen
gas into usable ammonium.
|
|
Euplotes
vannus
(Ciliate protozoan)
|
Polynucleobacter
necessarius
(Betaproteobacterium)
|
Obligate Endosymbiosis
(Survival dependency)
|
Metabolic Maintenance: The ciliate cannot survive
or divide without the cytoplasmic bacterium, which manufactures essential
cytoplasmic compounds.
|
Details on the Natures of
these of Key Species.
1. Mixotricha paradoxa
(The Ultimate Microbial Team)
Found inside the gut of the
primitive Australian termite Mastotermes darwiniensis, Mixotricha
paradoxa is famously known as the “poster organism” for symbiosis.
- Nature: The eukaryote itself is
physically incapable of moving on its own or producing energy through
oxygen. It serves as a structural chassis.
- Coalition: Roughly 250,000 helical Treponema
bacteria anchor themselves to specific structural brackets on the host’s
cell membrane. They
synchronous-wave like microscopic oars to propel the eukaryote forward. Inside the cell, spherical Endomicrobiellum
bacterial prokaryotes act as functional ‘mitochondria homologues,’
processing cellulose and generating cellular energy (ATP).
2. Paulinella
chromatophora (Evolution
Caught in the Act)
This freshwater amoebic
eukaryote represents the only known modern instance of a second independent
primary endosymbiosis. (The first instance occurred over ~1.5 billion years ago, since giving rise to all green
plants).
- Nature: Millions of years ago, a
predatory ancestor of the Paulinella
eukaryotes ingested a photosynthetic cyanobacterial prokaryote, but failed
to digest it. Instead, the two,
disparate units integrated permanently.
- Coalition: The cyanobacterial prokaryotic partner
(now a specialized structure called a “chromatophore”) has undergone
massive genome reduction, offloading its vital survival genes directly
into the eukaryotic amoeba’s nuclear DNA.
The single-celled amoeba controls the reproduction of the bacteria
so precisely that, when the eukaryotic host cell divides, the prokaryotes
divide with it, evenly.
3. Epithemia
Diatoms and their “Spheroidal Bodies”
Diatoms
are unicellular eukaryotic algae enclosed in elegant silica shells.
- Nature: Members of the eukaryotic family
Rhopalodiaceae
harbor modified cyanobacterial prokaryotes that look like small spheres
inside these eukaryotes’ cytoplasms.
- Coalition: Unlike standard plants, which
can photosynthesize but cannot pull nitrogen straight from the air, this
single-celled coalition does both. The prokaryotic spheroidal body has
completely lost the genes required to photosynthesize independently,
rendering it non-viable outside of the eukaryotic host. In exchange for
safety and food from the host, it acts as a highly efficient internal Nitrogen-fixation
fertilizer factory.
On the Dialectical, Aufheben Nature of the Opposition Between
Eukaryotic Cell units and Prokaryotic Cell units, constituting their categorial mutual opposition –
qe ~ qp.
The dialectical opposition
between eukaryotic single-cell units and prokaryotic single-cell units is one
that we cannot directly, sensuously perceive, since both are far too minute for
our unaided eyes, or fingers, to see or feel.
That opposition is produced via
a ‘concrete negation operation’, yielding a ‘‘‘negativity’’’, a specific
‘not-ness’, that distinguishes the two.
That [self-]negation
operation is not any “abstract negation”
operation, that obliterates its operands into nothingness [cf. Hegel].
It is a “determinate
negation” operation, a self- «aufheben»
self-operation, by which, e.g., multiple prokaryotic single-cell units coalesce
themselves into single eukaryotic single-meta-cell units, negating
their ‘prokaryoticity’ determination, but also conserving
that ‘prokaryote-ness’ inside their collective new ‘eukaryote-ness’, and uplifting
themselves into the higher size/mass/complexity scale of eukaryotic single-meta-cell
units, leaving behind their former ‘prokaryote-ness’-only complexity scale and
level.
These prokaryotic single-cell units thus collectively continue,
by adding a new level to, the multi-level ‘qualo-fractal’ scales-progression
that began, per our hypothesis, with/from the ‘spandetron’ units
of “Dark Energy” – of “empty” space itself – themselves.
Dialectical, Ontological Category-Symbols’
subscript Commutability and Meaning-Duality.
Note that category-symbol qep [generically q96 = q64+32] has the same ‘‘‘order-number’’’ [the same ‘ordinal-quality
generic subscript index’] as does category-symbol qpe [q96 = q32+64] – i.e., is the 96th category-symbol in this ‘Dialectic of Nature’,
‘cosmo-ontological’ category-symbols progression.
The latter, qpe, can be solved as representing another syntactically possible
‘[cosmo-]ontological’ process within prokaryotic cell units’/eukaryotic cell units’
combinations/complex unities/uni-categories/dialectical partial syntheses.
Specifically, subscripts-commutated
category-symbol qpe might well, for
example, be interpreted as describing the conversions
– ‘‘‘catalyzed’’’ by remaining prokaryotic cell units, of eukaryotic cell units’
biomass back down into prokaryotic
cell units’ biomass.
Category-Symbol qpe – Some Rather “Grisley”
Instantiations.
Not only does category-symbol qep have
instantiations, as seen above – both symbiotic examples,
qep <---> qpe,
and predatory examples, the latter as ‘conversion
formations’ that convert p ‘onto-mass’ up into e ‘onto-mass’; p ^ e.
Category qpe also has
instantiations, albeit rarer ones, on our planet, in the form of ‘conversion
formations’ that convert e ‘onto-mass’ back down into p ‘onto-mass’; e ¯ p;
qep ~ qpe.
Several such cases of the
latter kind of predation are described below.
While the overwhelming
ecological norm is for larger, single-celled eukaryotes (like amoebae
and ciliates) to hunt and engulf smaller prokaryote cellular units – archaeal cell
units and, especially, bacterial cell units – there are several remarkable,
documented cases where prokaryotes act as the predators and “eat”,
or ‘‘‘convert’’’, eukaryote single-cell units’ biomass
back down into [likewise single-cell unit] prokaryote biomass, by
‘‘‘consuming’’’ eukaryote cell units’ biomass.
Because prokaryotes generally
lack the machinery for phagocytosis (the ability to wrap a flexible membrane
around large prey to engulf it), they utilize specialized, alternative
predatory strategies.
The most notable and heavily
studied cases of single-celled prokaryotes preying upon single-celled
eukaryotes include:
1. The “Vampire” Bacteria (Vampirovibrio
chlorellavorus)
- The Eukaryotic Prey: Chlorella
units (a genus of single-celled green algae).
- The Mechanism (Epibiotic Predation): Vampirovibrio
attaches itself directly to the outer cell wall of a eukaryotic algae cell
unit. Instead of entering the cell,
it forms a specialized cytoskeletal protrusion (metaphorically, called a “fang”),
that pierces the eukaryotic cell unit’s cell membrane. The prokaryotic cell unit then releases
hydrolytic enzymes into the eukaryote, to liquefy the algal eukaryote’s insides,
and then literally sucks out the “food” that is the eukaryotic cell unit’s
cytoplasm, leaving behind an empty, dead shell of the former eukaryotic living
cell unit.
2. Ixotrophic Bacteria and
Contractile Injection Systems
- The Predator: Various ixotrophic bacterial prokaryotics cell units (e.g., from certain
marine strains).
- The Eukaryotic Prey: Diatoms
(single-celled eukaryotic algae units with silica shells).
- The Mechanism (Lytic Capture): "Ixotrophy" functions much like a “microscopic
flypaper”. The bacterial units catch passing eukaryotic cell prey units by
sticking them to the prokaryotic cell unit’s cell surfaces. Recent
research published in Science shows that these bacteria utilize Contractile
Injection Systems (CISs)—nanoscopic macromolecular daggers
structurally similar to bacteriophage tails—to punch holes in their
eukaryotic cell unit prey, inducing cell lysis (bursting) so the bacteria’
prokaryotic cell units can feed on the released eukaryotic prey nutritional
biomass.
3. Pack-Hunting
Myxobacteria (Myxococcus
xanthus)
- The Eukaryotic Prey: Single-celled fungi units (yeasts like Saccharomyces
cerevisiae) and various eukaryotic microalgae units.
- The Mechanism (Wolfpack Predation): While a single M.
xanthus prokaryotic cell unit is too small to take down a eukaryote
cell unit alone, these prokaryote units coordinate into massive, swarming “wolfpacks”. When the swarm encounters a population
of single-celled eukaryote cell units, they collectively secrete a potent
cocktail of lytic enzymes and antimicrobial peptides. This dissolves the
sturdy cell walls of the eukaryotic yeast or algae units from their
outsides, killing those eukaryotic cell units, and allowing the prokaryotic
cell bacterial units to swarm and absorb the eukaryotic cell units’ former
contents – their thus-extruded nutrient-rich runoff.
4. Endosymbiotic Prokaryotic
Units as Parasites, Acting
as Predators (Chlamydiae)
- The Predator: Primitive environmental chlamydiae prokaryotic cell units (e.g., Waddlia,
Parachlamydia).
- The Eukaryotic Prey: Acanthamoeba
(free-living, single-celled amoebae eukaryotic cell units).
- The Mechanism (Intracellular
Predation): Unlike medical
chlamydia, these environmental prokaryote single cell units target wild,
eukaryotic protist cell units. They exploit the amoeba’s natural tendency
to swallow bacteria. Once inside,
instead of being digested, the bacteria’ prokaryotic cell units breach the
digestive vacuole, replicate wildly within the eukaryotic host cell’s
cytoplasm, consume its energy reserves (ATP), and eventually rupture the eukaryotic
cell unit from the inside out, the resulting expanded population of these
prokaryotic single cell units to find new prey thus freed, virus-like,
to search out yet-new eukaryote cell unit prey.
Summary of Strategies
Because prokaryote predator cell
units are highly restricted by size, they bypass the more usual “swallowing” of
their prey, instead relying on:
- Epibiotic sucking: Latching on and draining their eukaryotic single
cell unit prey of its nutritive content.
- Chemical lysis: Injecting or secreting enzymes to force their eukaryotic single
cell unit prey to burst, releasing their eukaryotic single cell unit prey
of its nutritive content externally to its former cell unit confinement,
thus making it externally available to these predatory prokaryotic cell
units.
- Intracellular hijacking: The, much smaller, prokaryotic single cell units
entering into the inside of their, much larger, eukaryotic single cell unit
prey, to eat out their prey’s nutritive content from inside the eukaryotic
cell prey unit, from its inside, out.
The sample of specimens noted above show that the ‘dialectics-of-nature’,
combinatoric ontological-category symbol(s), qep, and qpe, ha(s)(ve) Earth-immanent instances that represent both predatory combinations and symbiotic
combinations of the e and p onto-types.
For more
information regarding these
Seldonian insights, and to read and/or download, free
of charge, PDFs and/or JPGs of Foundation books, other texts, and images, please see:
www.dialectics.info
and
https://independent.academia.edu/KarlSeldon
For partially pictographical, ‘poster-ized’ visualizations of many of these Seldonian insights -- specimens of ‘dialectical art’ – as well as dialectically-illustrated books
published by
the F.E.D. Press, see –
https://www.etsy.com/shop/DialecticsMATH
¡ENJOY!
Regards,
Miguel
Detonacciones,
Voting Member, Foundation Encyclopedia Dialectica [F.E.D.];
Elected Member, F.E.D. General Council;
Participant, F.E.D. Special Council for Public Liaison;
Officer, F.E.D. Office of Public Liaison.
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your comments on this blog-entry below!